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Borstein trained in improv at the ACME Comedy Theater, where she met her writing partner and future husband Jackson Douglas.
Borstein was also a writer and voice actor for several television shows, including Casper, Pinky and the Brain, and Power Rangers Zeo, before joining the cast of MADtv in 1997 as a featured player, becoming a repertory player mid-season.Borstein later starred as Dawn Forchette on the HBO comedy series Getting On, which is a remake of a UK series of the same name.The series has gained positive reviews and earned multiple Primetime Emmy Award nominations. In 2017, Borstein began starring as Susie Myerson in the historical comedy-drama series The Marvelous Mrs.In its second season, she held the position of writer and consulting producer.She also guest-starred as Lou Deckner in numerous episodes of the series, beginning in the first season episode "But at Last Came a Knock." She also appeared as a comedian contestant on IFC's short-lived comedy-driven game show Bunk in 2012.